The xylem is formed in the growing season by the cambium; in temperate regions the cells formed in the spring are larger in diameter than those formed in the summer, and this results in the annual rings observable in cross section. The new cells lose their protoplasm as they form the various tissues; the older, nonfunctional cells become plugged up, darken in color, and often accumulate bitter or poisonous substances (tannins, dyes, resins, and gums). This inner wood (the heartwood, as opposed to the functional sapwood) is valued for outdoor construction because of its resistance to moisture and to decay-producing organisms.
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